Thank you very much for the wishes, Svinleshas, much appreciated. This is the first chance I’ve had to post since my last message, I tried to post this response yesterday about a dozen times but the board was refusing to cooperate. Anyway, I pick up as I may from your list of examples before (hopefully) proceeding as time allows. This follows from my previous post, which I had to cut short.
The example of poor/good adaptation in this case was the male peacock, but I submit that that is poor adaptation only as far as an individual male peacock’s life is concerned. In genetic terms, if sexual advantages outweigh survival advantages then very gaudy displays may be selected for, because they will be reproduced more than the inferior displays of less-endowed competitors who are less successful at attracting mates (even if they live longer). After all, it’s the transmission of genes that is relevant to the evolution of the peacock, not necessarily the long-term survival of individual males but how successfully they are able to reproduce and therefore spread their genes. Consider that a long, colourful tail and train signal not just beauty but also a peacock’s health: firstly, because bright, vibrant plumage in birds are signals of good health and relatively low parasitic activity; secondly, a male who manages to survive and thrive in spite of the impediment of a long, visible tail must be exceptionally fit for survival, and therefore good breeding material. Thus females’ attraction to these visually exorbitant characteristics would select for the characteristics, a situation that, overall, strikes me as quite advantageous for the peacock species.
But when you look at a morelet crocodile, it has adapted to deal with colder water than its relatives live in. You therefore wouldn’t expect to find a crocodile of the variety usually confined to warm waters in the relatively cold environment of morelets, and as far as I know you don’t (falsifiable prediction).
The above (including and especially the peacock) are examples of the levels of complexities involved in the study of organisms. Look superficially at just one or two features of a creature, and you could easily come to conclusions that are superficially contradicting. (or unfalsifiable).
No of course not, but the general observation is that organisms are well adapted to their environments and that those not well adapted either adapt or die off/fail to compete effectively; these assertions, I hope we have seen, are easily falsifiable. Specific observations, such as the cunning design and placement of the crocodile’s eyes and nostrils, or the thumblessness of certain primates, are subject to different and variable criteria because such adaptations necessarily serve different purposes and result from different, highly variable conditions (plus, they may not be immediately clear without a detailed look at the organism in question and its niche). In the case of the African and other colobus, the loss of the thumb actually seems to be an advantage to the creature: not only is there no misaligned digit to get snagged on a branch during the colobus’s regular high-speed acrobatics, not only is the hand grip on branches stronger as a result, but the monkey doesn’t seem to miss its thumb terribly much, manipulating objects by using fingers against palm instead of the more widespread primate manipulation strategy of the opposable thumb.
What’s interesting is that species of colobus are among the most arboreal of primates, seldom even setting foot on the ground, and are among the best adapted to arboreal life of all primates. It would seem that the colobo’s loss of thumbs allows it to move faster and more efficiently in the tree canopy – certainly in this case the logic is perfectly compatible with the observation - but the matter is difficult to falsify immediately without more data (it doesn’t mean it’s not possible, of course; I suppose an extensive study of the evolutionary advantages of thumb versus thumbless primate arboreal locomotion may help confirm that).
I discussed a number of the crocodile’s adaptations in the last thread, and noted how each feature we looked at (nictitating eye membrane, nostrils and eyes just above water level allow it to camouflage itself, etc.) is well suited to the crocodile’s preferred environment. The concept of natural selection is falsifiable in this case because we could (conceivably) find poorly adapted crocodiles competing with other well adapted crocodiles in the latter’s habitat (if we did, we would not observe them to last very long, but it’s conceivable they might and that should meet the falsification criterion). That is what my example of a crocodile with eyes and nostrils on the underside of its head was supposed to illustrate: such a poorly adapted croc could see and smell very little of its prey, and it couldn’t even breathe in water while lounging lazily around as crocs are wont to do waiting for meals. I consider this example a funny little thought experiment, certainly not the height of scientific inquiry but illustrative of the problem I named so clumsily.
Perhaps a biologist could help us out here. The short version is that the human genome should provide a record of our evolution. Common descent, a theory that goes hand-in-hand with evolution, postulates that the earth’s biota are all genealogically related, and differences are due to varying evolutionary paths. We can therefore predict the existence of substantial similarities in the genetic record across species that are taxonomically close, and somewhat less substantial similarities in more distant species. This brand new study is particularly interesting because it lends weight to a hypothesis some observers had regarding our shrew-like ancestors and our phylogenetic status quo:
*from the press release at http://www.genome.gov/11008356 *
Pioneering Study Compares 13 Vertebrate Genomes
Multi-Species Approach Provides Unprecedented Glimpse Into Function and Evolution of the Human GenomeBy systematically comparing the patterns of a certain type of genomic change, called transposon insertions, among the different species’ sequences, these investigators were able to address a heated controversy in the field of evolutionary genomics. Their analyses confirm recently proposed trees of mammalian evolution indicating that primates (human, chimpanzee, baboon) are more closely related to rodents (mouse, rat) than to carnivores (cat, dog) or artiodactyls (cow, pig). Indeed, the evidence revealed by the new sequence data refutes alternative evolutionary trees that place rodents much farther away from primates.
A logical, evidence-supported, and falsifiable explanation to account for the numerous protein domains that humans share with other species is evolution from common descent. For a more hardcore biological approach and supporting data see W.H. Li et al., Evolutionary analyses of the human genome, Nature, February 15, 2001.
Now let’s look at the idea of common descent a bit more closely. From a link already provided:
According to the theory of common descent, modern living organisms, with all their incredible differences, are the progeny of one single species in the distant past. In spite of the extensive variation of form and function among organisms, several fundamental criteria characterize all life. Some of the macroscopic properties that characterize all of life are (1) replication, (2) information flow in continuity of kind, (3) catalysis, and (4) energy utilization (metabolism). At a very minimum, these four functions are required to generate a physical historical process that can be described by a phylogenetic tree.
If every living species descended from an original species that had these four obligate functions, then all living species today should necessarily have these functions (a somewhat trivial conclusion). Most importantly, however, all modern species should have inherited the structures that perform these functions. Thus, a basic prediction of the genealogical relatedness of all life, combined with the constraint of gradualism, is that organisms should be very similar in the particular mechanisms and structures that execute these four basic life processes.
Confirmation:
The structures that all known organisms use to perform these four basic processes are all quite similar, in spite of the odds. All known living things use polymers to perform these four basic functions. Organic chemists have synthesized hundreds of different polymers, yet the only ones used by life, irrespective of species, are polynucleotides, polypeptides, and polysaccharides. Regardless of the species, the DNA, RNA and proteins used in known living systems all have the same chirality, even though there are at least two chemically equivalent choices of chirality for each of these molecules. For example, RNA has four chiral centers in its ribose ring, which means that it has 16 possible stereoisomers - but only one of these stereoisomers is found in the RNA of known living organisms.<snip>
Potential Falsification
Thousands of new species are discovered yearly, and new DNA and protein sequences are determined daily from previously unexamined species (Wilson 1992, Ch. 8). At the current rate, which is increasing exponentially, over 41,000 new sequences are deposited at GenBank every day, amounting to over 40 million new bases sequenced every day. Each and every one is a test of the theory of common descent. Based solely on the theory of common descent and the genetics of known organisms, we strongly predict that we will never find any modern species from known phyla on this Earth with a foreign, non-nucleic acid genetic material. We also make the strong prediction that all newly discovered species that belong to the known phyla will use the “standard genetic code” or a close derivative thereof. For example, according to the theory, none of the thousands of new and previously unknown insects that are constantly being discovered in the Brazilian rainforest will have non-nucleic acid genomes. Nor will these yet undiscovered species of insects have genetic codes which are not close derivatives of the standard genetic code. In the absence of the theory of common descent, it is quite possible that every species could have a very different genetic code, specific to it only, since there are 1.4 x 10[sup]70[/sup] informationally equivalent genetic codes, all of which use the same codons and amino acids as the standard genetic code (Yockey 1992). This possibility could be extremely useful for organisms, as it would preclude interspecific viral infections; however, it has not been observed, and the theory of common descent effectively prohibits such an observation.
That last paragraph ought to be of particular interest. I’ll let my betters in this camp discuss the probabilistic nature of evolutionary science. I think that may be an unnecessary refinement for the purposes of this conversation, however I recall it was brought up the first time a couple pages back.