First of all, the US Coast Guard is not normally a part of the Department of Defense (although it can be transferred to the Department of the Navy in wartime). It was formerly a branch of the Department of Transportation, and now belongs to the Department of Homeland Security (where it utterly doesn’t belong, but whatever).
The elite operations unit in the USCG are the Aviation Survival Technicians, commonly referred to as “rescue swimmers” and publicized in the (execrable) Kevin Costner-Aston Kutcher film The Guardian. The Air Force SpecOps units include the Air Force Pararesuce Jumpers (PJs), Weather Operators, Tactical Air Command and Control Specialists, and the operators of the 160th Special Operations Aviation Regiment, which performs infil/exfil for the Army and other branches as necessary. The Central Intelligence Agency also has its own covert operations arm called the Special Activities Division, largely recruited (at least historically) from the Marine Corps Division Recon and Force Recon.
IMHO, PJs are the most bad-assed of the bad. Most SpecOps go into a hot zone on the prowl, perform reconnaissance, sabotage, hostage rescue, whatever, and then blow out hot to the r.v. On the other hand, PJs go into a hot zone, hunt around for a downed flier or injured soldier/marine, and then fight their way out. Purportedly, these guys never have to buy a drink in any military-favored bar as any soldier, sailor, flyer, or marine will be more than happy to pay the tab.
In general, the tensions between branches are attenuated in the Special Operations community, partially because of the deliberate effort to unify command, but also because unlike traditional units in the different branches, these guys all have pretty common missions and can’t allow service rivalry to cause friction. This also means that it is fairly easy to shift from one service to another; a lot of Delta operators come from the Navy UDT, Marine DevRecon, or even non-military sources, as well as from normal Ranger units. Because there are both high athletic and intellectual standards for entry and acceptance into the Special Operations units, they tend to be both less picky about the provenance of candidates, and more permissive about behavior (within a certain range) than traditional units. That being said, you can’t just walk in off the street and demand that a recruiter put you up for selection; you basically have to be nominated by someone within or attached to the community to even get the invite (typically after at least two tours or equivalent experience) and then have to pass a grueling and intentionally demoralizing selection process before even entering the training regime.
Stranger