Ungulates in which the weight is borne on two main hooves are known as the even-toed ungulates, orartiodactyls. As has been said, their main toes are the third and fourth. All the artiodactyls have lost the equivalent of the thumb (digit I). Many have small dewclaws behind the main hooves, representing the second and fifth toes, but some like camels have lost them.
Horses, rhinos, and tapirs are odd-toed ungulates, or perissodactyls, in which the weight is borne on the middle toe. Horses have lost all the other toes. Tapirs have four toes on the front feet (II, III, IV, V), and three on the rear (II, III, IV). Rhinos have three toes on each foot (II, III, IV).