Usain Bolt (and every other sprinter) does not lean forward when sprinting at top speed. (The do during the acceleration phase when raising their COG from the crouch position. If they stood up immediately, they would be moving their COG backwards, and basically be stationary for the first half-second or so. A standing start is faster for the first 20-35 metres of the race).
The hardest part of raising the COG is the head - this is a 5-6 kgs lump sitting (wobbling) on top of a set of relatively weak stabilising muscles (neck), with no muscles of its own to assist in the running motion, transfer of weight or balance. Most people find a comfortable position when walking or running (or even standing) is to hold the head slightly forward of the COG - this leads to the impression that runners ‘lean forward when running’.
I you watch any top sprinter on Youtube you will notice that they are nearly exactly upright once they reach maximum or near-maximum speed. The foot strikes the ground slightly in front of the body, and applies force only while the foot is in front of the body. The keys to maximising sprint top speed are:
- Applying maximum force to the ground each stride (how hard you can push off)
- Applying that force quickly (how fast you can push off)
Using these basic ideas you need to be a) strong and b) have quick-acting muscles - the fast-twitch muscles, gene ACTN3 etc. The sprint action that top athletes use is a learned action quite a bit different to the ‘natural’ fast running style we all use from the age of 3.
Long legs have pros and cons - longer strides, but also longer cycle times. The one definite advantage they do have is that fewer strides mean fewer strides to fatigue the body - meaning you can hold maximum speed for longer (it’s a slight advantage, but it seems to be confirmed by the studies - yes there are factors like longer legs are heavier and take more effort etc, but overall - better to have long legs. The advantage is very slight).
Carl Lewis was 3 inches shorter but 5-6 KGs heavier than Usain Bolt. Ben Johnson was about 4 inches shorter than Lewis, but probably the same weight, or even heavier, and significantly stronger (illegally). He was faster, too.
It’s all a case of balancing out the pros and cons of the various factors - good muscles mean more force, but more weight. Too much muscle mass restricts speed of and range of movement. Flexibility is important - too much flexibility can mean less stability in applying force to the ground. Etc, etc.
Top sprinters are freaks of nature, who have been trained. Usain Bolt is a well-trained freak among freaks.