Consider this…If higher order animals had wheels, the only advantage would be when gravity is working with you (i.e. downhill). It would seem like it isn’t very energy conservative. When you walk uphill, or even level for that matter…your energy expended by taking a step and placing it on a surface, moving your leg and placing it on a surface maybe 2 or 3 feet in front of you. A wheel has constant contact with the surface it is on. Since contact equals friction, friction equals resistance, and resistance requires a greater energy expenditure to complete a task, it would seem that a wheel would be more of a hindrance then a benefit. Besides, wheels wear down…when was the last time someone had to have the soles on their feet replaced?
This is in fact a comment on Cecil’s column: Why has no animal species ever evolved wheels?, so I have moved the topic to the appropriate forum. – CKDextHavn